Sunday, April 28, 2013

Temple Flat Rock (Wendell, Wake County, North Carolina)

Triangle Land Conservancy's Temple Flat Rock Preserve

Outside Wendell, NC (pop. ~ 6,000), an eastern satellite town of Raleigh, sits 5,270 square meters of exposed granite rock that supports a unique community of lichens, bryophytes, and angiosperms (flowering-plants). In 1984, the Temple family donated this unusual Registered Natural Heritage Site to the Nature Conservancy. In the mid-1990s, Temple Flat Rock was transferred the Triangle Land Conservancy (TLC), becoming the organization's first conservation easement.

Temple Flat Rock, Wendell NC with Appalachian sandwort (white flowers) and elf-orpine (pink); April 2013


Fence lizard at Temple Flat Rock
Temple Flat Rock, a granite outcrop that showcases a large expanse of the Rolesville granitic batholith, features a number of endemic plants, including Appalachian sandwort (Minuartia glabra) and elf-orpine (Diamorpha smallii). It also supports a number of mosses, over 44 species of lichen, hardy eastern-red cedars, and a few fence lizards and ground skinks that use Temple Flat Rock to bask in the sun on cool days.

Piedmont prairie establishment at Temple Flat Rock




The TLC preserve is also notable as a Piedmont prairie restoration site. Like the tallgrass prairies of the Midwest or the longleaf pine forests of North Carolina’s coast, the savanna-prairie complex of the Piedmont is a nearly extinct ecosystem. Today, prairie restoration sites occur around Charlotte and the Triangle. Yet, few of these places adequately capture the essence of the extirpated Piedmont savanna: an expanse of tussocky tall grasses and sky-reaching forbs beneath an open canopy of hardy post oaks, sometimes stretching for miles before meeting a riparian forest or wooded hill. These ecosystems, described by nearly every European explorer that had visited North Carolina’s Piedmont between the mid-16th and mid-18th centuries, were nearly forgotten landscapes even by the start of the American Civil War. Fortunately, Walt Tysinger, the land manager at TLC, has been working hard to bring these systems back both at Horton Grove (by Stagville plantation, north of Durham, NC) and at Temple Flat Rock.

Currently, Temple Flat Rock includes about 5 acres of granite outcrop, 15 acres of mixed hardwoods, and 16 acres of grassland established from old agriculture fields and horse pasture. Like most prairie restoration, recreation, and establishment managers, Tysinger struggles to control the non-native fescue (Festuca sp.), sericea lespedeza (Lespedeza cuneata) and native sweetgum (Liquidambar styraciflua). Using a combination of dormant season burns, spot spraying of herbicides, and bush hogging, Tysinger has been able to control the grassland invasives enough to establish (from plugs) native warm-season grasses, such as Indian grass (Sorghastrum nutans), little blue stem (Schizacharyium scoparium), and splitbeard bluestem (Andropogon tenarius). Other species seen in the grassland establishment include prickly pear (Opuntia humifusa), common milkweed (Asclepias syriaca), and broomsedge (Andropogon virginicus).

Chickasaw plum (Prunus angustifolia) thicket in Temple Flat Rock grassland establishment.


Prickly pear (Opuntia humifusa) at Temple Flat Rock grassland.


In the Triangle, a number of remnant grasslands or restoration projects occur on the basic soils of the Triassic Basin, underlain by magnesium- and iron-rich diabase. These soils often contain montmorillonite, a type of clay that shrinks and swells so dramatically that it can deter tree root growth. In contrast, the Temple Flat Rock prairie establishment project occurs on acidic Louisburg (Typic Hapludults) and Appling (Typic Kanhapludults) soil. Yet, the site is so xeric and well-drained, that with a little ecological disturbance (e.g., fire), a grassland ecosystem feels perfectly natural. Moreover, the hard work of Tysinger and the Triangle Land Conservancy serves to remind us all of the natural and cultural heritage of North Carolina's Piedmont prairie landscape.

Throwdown from Hurricane Fran in the oak-hickory forest at Temple Flat Rock.

For more information about Temple Flat Rock, please visit the TLC website at www.triangleland.org.



Monday, April 15, 2013

West Point on the Eno (Durham, NC): Emerging Dragonfly and Spring Flora II

West Point on the Eno: Emerging Dragonfly & Mid-Spring Flora II

Emerging Dragonfly: on the bridge by the mill

Rue anemone (Anemonella thalictroides): north facing hillside, near Sennett's hole

Pinxter-flower (Rhododendron periclymenoides)





Sunday, April 14, 2013

West Point on the Eno (Durham, NC)

West Point on the Eno (Durham, NC) -- Mid-Spring Flora, Basking Snakes, and Unidentified Crayfish

Creeping Charlie (Glechoma hederacea): found in the bottomlands near Sennett's Hole;
weedy, invasive ground cover from Europe; toxic to cattle and horses, but used medicinally. 

Golden Alexanders (Zizia aurea): found in the bottomlands near Sennett's Hole

Spring beauty (Claytonia virginica): found in the bottomlands near Sennett's Hole

Early saxifrage (Saxifraga virginiensis) -back of flower head, found near "turtle rock" of rocky bluffs at West Point on the Eno

Early saxifrage (Saxifraga virginiensis) -with distinctive basal leaves, found near "turtle rock" of rocky bluffs at West Point on the Eno

Northern water snake (Nerodia sipedon): adjacent to the mill wheel at West Point on the Eno

Queen snake (Regina septemvittata): adjacent to the mill wheel at West Point on the Eno

Crayfish, spp. unknown: Warren Creek
for more information on North Carolina crayfish species, please visit: http://216.27.39.101/wildlife_species_con/nccrayfishes/nc_crayfishes.html

Saturday, April 13, 2013

Return to Mason Farm (Chapel Hill, NC)

Redbelly Water Snake (Nerodia erythrogaster) at Mason Farm, Chapel Hill, NC


One evening this April, I visited Mason Farm with my dear friend gumby. As we started hiking down the grassy path, I commented on how Mason Farm was always a lucky place to find snakes, especially brown snakes, rough green snakes, and rat snakes.

We were lucky to add another species to that list: redbelly water snake (Nerodia erythrogaster). This juvenile rested in the middle of the path in a grassy field adjacent to wetland habitat. I've yet to find a red-bellied water snake in the water, or even on a log sticking out of the water. I often find them sunning themselves in an open grassy area, near but relatively removed from water.

Redbelly water snakes have been observed at Mason Farm before, but there is still something exciting about seeing a species at the edge of its range. In North Carolina, redbelly water snakes are more abundant in the Coastal Plain, and their distribution only sneaks into the eastern Piedmont counties: Richmond, Lee, Moore, Wake, Chatham, Orange, and possibly Anson.


Sunday, November 11, 2012

Mason Farm Biological Reserve, Chapel Hill, NC

Brown Snakes at Mason Farm Biological Reserve (Chapel Hill, NC)

One of the most delightful surprises of warm autumn days is discovering snakes warming up on walking paths through forest and field. Today temperatures reached at least 72°F in the Triangle, and snakes were out basking in the sun. At Mason Farm Biological Reserve, we discovered three different brown snakes (Storeria dekayi) warming up on the graveled path. Brown snakes are recorded year round in North Carolina, even as late as December 23 in Guilford County. 





References:
Palmer, W. M. and A. L. Braswell. 1995. Reptiles of North Carolina. University of North Carolina Press: Chapel Hill, NC.

Sunday, September 9, 2012

GREAT EXPECTATIONS: September in the Piedmont


ON THE WILD SIDE

Birds.− September brings a number of winter residents back to the Piedmont, including the yellow-bellied sapsuckers, an occasional red-breasted nuthatch and a number of wrens (winter, sedge, marsh) and sparrows (swamp and white-throated). Ruby-crowned kinglets, tiny olive-grey birds with bright red spots on their crown, returned to the Piedmont this month after spending the summer in the coniferous forests of Canada and the northern United States. Migrant warblers continue to pass through North Carolina as they head towards the neotropics for the winter. Birders may even see snowy egrets, little blue herons and tricolored herons, which won’t return again to the Piedmont until early April.

Did you know that not all birds need to drink water? Hummingbirds rely on nectar to stay hydrated. Birds in arid areas may not drink at all either (think: ostriches.) Yet, most birds do drink to replenish fluids lost by breathing, excretion through skin and waste production. Some submerge their bills into the water and simply suck it up (e.g., doves). Other birds dip their bills into the water and then point up to the sky, letting the water fall back into their throat. A number of small bird species drink dew-drops.

Butterflies.− This time of year, butterflies are often surprisingly abundant. Look out for the usual suspects, including hackberry emperors gleaning sap from trees, tiger swallowtails puddling to uptake salts and other nutrients, and pearl crescents, whose caterpillars feed almost exclusively on asters.

Expect to see a pulse of cloudless sulphurs, little yellows and sleepy oranges this month. Swallowtail sightings will likely drop-off by mid-month, with the exception of the black swallowtail. Butterfly watchers can also expect to see gray and red-banded hairstreaks, gulf and variegated fritillaries, as well as an increased number of viceroy sightings. Monarchs may be seen as they migrate southward to their winter residence in Mexico.

The Carolinas are home to five families of butterflies: the skippers (Hesperiidae), gossamer wings (Lycaenidae), brush-foots (Nymphalidae), swallowtails (Papilionidae) and the sulphurs and whites (Pieridae). Each of these families can be divided into a number of sub-families, each having distinct identifying characteristics.

This month, we will consider the hairstreaks (subfamily: Theclinae), small and intricately patterned members of the gossamer wing family (Lycaenidae). Hairstreaks are named for the small hair-like tails on the end of each hindwing. These tails resemble antennae and, along with bright eyespots, trick predators into attacking the tips of the wings, rather than the soft body of the butterfly. Males and females usually differ in appearance (i.e., they are sexually dimorphic), but both sexes fly erratically and perch with their wings held together while moving their hindwings up and down.  Gray hairstreaks (Strymon melinus), the most widely and commonly seen hairstreak in North Carolina, is particularly abundant in September. They are blue-gray below, with bright orange spots and a dark tail with a white tip. Gray hairstreaks prefer open sites, and larvae feed on partridge pea, vetch, clovers and other legumes.

Reptiles & Amphibians.− Continue to keep an eye out for snakes and turtles, especially baby box turtles. Skinks and toads are also out in abundance. Although you may still hear frogs and toads calling this month, large choruses won’t start up again until January.

Other Insects.− This month, expect an increase in praying mantis and spider activity. Praying mantises will exude their eggs in a frothy, hardened mass called an ootheca in September.  Meanwhile, female garden spiders (Argiope aurantia) will be guarding egg cases.

In Bloom this Month.− Be on the lookout for some striking September flowers:

In Bloom:
WINGSTEM – Actinomeris alternifolia
WHITE SNAKEROOT – Ageratina altissima
PARTRIDGE PEA – Chamaechrista fasciculata
TURTLEHEAD – Chelone glabra
BEECHDROPS – Epifagus virginiana
DEVIL'S-GRANDMOTHER - Elephantopus tomentosus
BONESET – Eupatorium perfoliatum
BLACK-EYED SUSAN – Rudbeckia sp.
TRAILING WILD BEAN - Strophostyles helvula
GREAT LOBELIA – Lobelia siphilitica
DOWNY LOBELIA – Lobelia puberula
SMALL SKULLCAP – Scutellaria parvula
AXILLARY GOLDENROD – Solidago caesia
GOLDENROD(S) – Solidago spp.

In Fruit:
PERSIMMON – Diospyros virginiana
HEARTS-A-BUSTIN’ – Euonymus americanus
MUSCADINES – Vitis rotundifolia

Wildlife Profile.− This month’s wildlife profile is the COPPERHEAD (Agkistrodon contortrix), the most widespread of North Carolina’s six venomous snakes. The copperhead probably occurs in every county in North Carolina, and is distributed throughout the southeastern United States. Known as “highland moccasins,” copperheads inhabit wooded areas ranging from riparian habitat to ridgetops. They sometimes reside in more open habitat and are fairly tolerant of human development, often frequenting trash piles and abandoned buildings.

The copperhead is a stout, moderately large viperid than can attain a maximum length of almost 4.5 feet. Adults are pinkish-brown with darker, brownish hourglass-shaped crossbands. Neonates, or newborns, have bright greenish-yellow tail tips. They mate in both spring and fall, and give birth to around a dozen live young in September and October. In autumn, copperheads will gather to den communally and with other snake species to better endure the colder months. Normally a quiet, retiring snake, copperheads will strike vigorously if annoyed.

Did you know?
  • Copperheads in the North Carolina Piedmont are intergradations of both northern copperheads (Agkistrodon contortrix mokasen) and southern copperheads (Agkistrodon contortrix contortrix).
  • Vipers are identified by their triangular heads and vertical pupils; Colubrids, non-venomous snakes, have circular pupils.
  • Copperheads in North Carolina eat cicadas, caterpillars, frogs, toads, birds, mice, shrews, voles, lizards, hatchling box turtles, ringneck snakes and worm snakes.

References:
National Audobon Society. 1980. Field Guide to North American Insects and Spiders. New York, New York: Chanticleer Press.
Conant, R., and J.T. Collins. 1991. A Field Guide to Reptiles and Amphibians: Eastern and Central North America. Houghton Mifflin Company: Boston, MA.
Cook, Dave. 2001. The Piedmont Almanac. Chapel Hill, North Carolina: Mystic Crow Publishing.
Daniels, J. C. 2003. Butterflies of the Carolinas. Cambridge, MN: Adventure Publications, Inc.
Ehrlich, P., D. Dobkin, and D. Wheye (1988). The Birder's Handbook. New York: Simon & Schuster.
LeGrand, H. E. Jr. 2009. Notes on the Butterflies of North Carolina. Available at: http://149.168.1.196/nbnc/
Palmer, W. M. and A. L. Braswell. 1995. Reptiles of North Carolina. Chapel Hill, NC: University of North Carolina Press.

Tuesday, July 31, 2012

Soil Ecology Class (starts Saturday!)


Interested in Soil Ecology? Sign up for a class at the North Carolina Botanical Garden. It will be a lot of fun!

Dates:Saturdays, Aug 4, 11, 18
Time: 1:30 - 5:30 pm
Instructor: Nicolette Cagle, Ecologist

This course is intended for a broad audience. Students are introduced to the complex world of soils including information on how they are formed, characterized, and populated by a wide array of organisms. An overview of soil types is presented, followed by the study of typical Piedmont soils and their properties. The various roles that soils play in both human society and ecological systems are discussed.

More information can be found here: http://ncbg.unc.edu/classes-workshops/#native_plant

Tuesday, June 19, 2012

2012 Call for Nominations for NAAEE Excellence in EE Awards


2012 Call for Nominations for NAAEE Excellence in EE Awards

Nominations Deadline: July 31, 2012
Help NAAEE recognize individuals and organizations that excel in EE by nominating them for one of our annual awards, including our highest honor -- The Walter E. Jeske Award.

The Call for Nominations is now open, and the awards will be presented on October 12th at the Annual Awards Luncheon at the 2012 NAAEE Annual Conference. Online nomination forms are linked here:

http://www.naaee.net/programs/awards

Monday, April 16, 2012

GREAT EXPECTATIONS: April in the Piedmont

Birds.− This month, spring migrants are headed our way. Some species that may be seen during the next couple months, as they head north, include: some wood-warblers (including golden-winged, Nashville, chestnut-sided, magnolia, black-throated blue, black-throated green, bay-breasted and blackpoll), herons (e.g., little blue heron, black-crowned night heron, cattle-egret), thrushes (e.g., veery, grey-cheeked and Swainson’s) and sandpipers (e.g., spotted and solitary). Also be on the lookout for sora, Virginia rail or a rare glossy ibis.

Species will be arriving this month with the intention of staying the summer and breeding here in the Piedmont; these include: whip-poor-wills, chimney swifts, ruby-throated hummingbirds, eastern wood-pewees, Acadian flycatchers, eastern kingbirds, northern parulas, prairie warblers, summer and scarlet tanagers, and yellow-breasted chats.

Some of our year-round residents are busy this month as well. Many Carolina wrens – small, energetic brown birds with upturned tails, distinct whitish eyebrows and curved bills -- hatch in April, and the young are heard boldly chirping in their nests. The female usually incubates five eggs in a nest of twigs, bark, leaves and grass busily constructed by both parents. Nest sites often can by found in cavities and protected areas, both natural and man-made. Unused grills and back porches are often prime real estate for these adaptable birds. After two weeks of incubation, young Carolina wrens hatch and noisily demand food from both parents. Carolina wrens live approximately six years and mate for life.

Did you know? The spring migration of birds occurs along four principal “flyways” in North America. Lucky for us, the Atlantic flyway crosses North Carolina, and provides a route northward from Central America and the West Indies for about 150 species of migratory birds. Ample food and cover exist along the entire mountain-free flyway, which stretches from the Gulf of Mexico, through the Carolinas and Virginia to the northeastern states and into central Canada.

Butterflies.− In April, butterfly watchers may begin to find some of the skippers (e.g., zabulon, dusted, pepper and salt), duskywings (mottled, zarucco), cloudywings (southern, northern, confused), satyrs (gemmed, Carolina) and pearlyeyes (southern and northern). Silvery checkerspots can be spotted in moist floodplains or sometimes near drier woodland borders, where adults glean nectar from, and caterpillars feed, on sunflowers (Helianthus) and rosinweeds (Silphium spp.). Look for red-spotted purples in hardwoods forests and forest edges; adults may be found taking sustenance from tree sap or damp ground, while caterpillars feed on cherries (Prunus spp.) and other members of the Rosaceae. Butterfly aficionados will continue to see a number of sulphurs and hairstreaks this month, as well as questionmarks and commas. The most spectacular visitors, this month, may be the monarchs and their mimics, viceroys.

Swallowtails, a largely tropical family of colorful butterflies with distinctive tails on their hind wings, make exciting sightings in April. Five swallowtail species make their homes in Durham County: the Spicebush Swallowtail, Pipevine Swallowtail, Black Swallowtail, Zebra Swallowtail, and the Eastern Tiger Swallowtail. The Eastern Tiger Swallowtail is also the state butterflies of Delaware, Georgia, South Carolina, and Virginia. Swallowtails typically use as a wide variety of flowers as nectar plants, and the caterpillars of some species can be quite particular. For example, the Pipevine Swallowtail caterpillars feed exclusively on Aristolochia species, including the native Dutchman's pipe (Aristolochia durior) and Virginia snakeroot (Aristolochia serpentaria). The Zebra Swallowtail caterpillars strictly rely on the two pawpaws species native to North Carolina, tall pawpaw (Asimina triloba) and dwarf pawpaw (Asimina parviflora) as foodplants.

Other Insects.− This month, field crickets will begin to call, crane flies will hover in the grass and ticks abound. Also expect to see some dragonflies zipping through the air, searching for mosquitoes and other prey. Dragonflies to look for in April include the darners, a family that represents some of the largest and fastest flying dragonflies in North America. Species sighted in Durham County include the Common Green Darner, Springtime Darner, and Swamp Darner. The Common Green Darner – a three inch long green dragonfly with a brown and yellow (females) or bluish (males) abdomen -- probably is active longer than any other Dragonfly species in the state: it can be seen in the Piedmont from March through October cruising over open habitat, especially near still water.

Reptiles & Amphibians.− This month, northern cricket frogs, eastern narrow-mouthed toads and Cope’s gray treefrogs will begin to call. American and Fowler’s toads, spring peepers, bull frogs, green frogs, southern leopard frogs and eastern spadefoots will continue to call, but the large choruses of southeastern chorus frogs will be winding down this month.

April frog call guide:

southeastern chorus frog: raspy, rising call like someone dragging their thumb over the teeth of a comb

spring peepers: a loud, medium pitched “peeep”

northern cricket frogs: clinking like two small metal balls being tapped together

American toads: long, musical trill

Fowler’s toads: long, slightly nasal, crabby trill

eastern narrow-mouth toads: buzzy and sheep-like call (like a Fowler’s toad, but shorter and buzzier)

eastern spadefoot toads: a crabby, deep “eeeerrrr”

Look in shallow permanent or ephemeral ponds to find frog and toad eggs and even tadpoles. In April, visible eggs include the long gray-green strings of Fowler’s toad eggs and globs of gelatinous black spotted Cope’s gray treefrog eggs. In most years, small and dark American toad tadpoles will emerge this month.

Also, be on the lookout for basking yellow-bellied sliders and painted turtles. Snakes will be out as well, so be sure not to step on the diminutive and well-camouflaged northern brown snake (Storeria dekayi) when walking on preserve trails. When gardening this month, also watch out for secretive rough earth snakes and eastern worm snakes, North Carolina’s most common snake species.

In Bloom this Month.− April is a wonderful month to test your tree identification skills. Try to identify trees by their bark or buds before they flower and leaf out!

In late March and early April, woodland hikers may notice a diminutive plant with three mottled leaves radiating out from the center, topped by a single maroon flower; this is likely one of the Piedmont’s most common trilliums, little sweet betsy or Trillium cuneatum. This musk-scented gem was once used medicinally to treat gangrene and skin ulcers. It is now a favorite among natural landscapers, as it is deer resistant.

In Bloom:

BUCKEYES (Aesculus spp.)

WINDFLOWER (Anemonella thalictroides)

SWEET-SHRUB (Calycanthus floridus)

MUSCLEWOOD (Carpinus caroliniana)

FLOWERING DOGWOOD (Cornus florida)

RATTLESNAKE-WEED (Hieracium venosum)

QUAKER-LADIES (Houstonia caerulea)

EASTERN YELLOW STAR-GRASS (Hypoxis hirsuta)

DWARF CRESTED IRIS (Iris cristata)

CORAL HONEYSUCKLE (Lonicera sempervirens)

HAIRY WOODRUSH (Luzula echinata)

VIRGINIA PENNYWORT (Obolaria virginiana)

MAY-APPLE (Podophyllum peltatum)

EARLY SAXIFRAGE (Saxifraga virgininiensis)

AMERICAN BLADDERNUT (Staphylea trifolia)

GIANT CHICKWEED (Stellaria pubera)

FOAMFLOWER (Tiarella cordifolia)

CATESBY’S TRILLIUM (Trillium catesbaei)

LITTLE SWEET BETSY (Trillium cuneatum)

MAPLE-LEAF VIBURNUM (Viburnum acerifolium)

DOWNY ARROW-WOOD (Viburnum rafinesquianum)

Soil Series of the Month.− Recall that the United States contains over 19,000 different soils series, i.e., the most specific grouping of soils based on shared history, chemistry, and physical properties. The most general classification of soil is the soil order. Worldwide, there are only 12 soil orders. North Carolina is home to seven: Entisols, Inceptisols, Alfisols, Ultisols, Mollisols, and Spodosols.

A common Ultisol, the quintessential deep red, clayey soil of the Piedmont, in this area is the Mayodan soil series. Mayodan soil is light gray or yellow-brown in the top six to twelve inches, and then becomes yellowish-red as depth increase. It is typically weathered from Triassic sediment (think: shales, sandstones, mudstones formed from erosion nearly 200 million years ago) in the uplands of the Piedmont and naturally supports oak-hickory forests. Over half of the acreage of Mayodan soil now support agricultural crops, especially corn, soybeans, tobacco, and cotton.

Wildlife Profile.− This month’s wildlife feature is the NORTHERN PARULA (Parula americana), a colorful warbler that migrates to the Piedmont in spring and breeds locally. Northern Parulas are small songbirds with blue-gray heads and wings, yellow throats, and a chest banded with black, red, and bright yellow. They also sport white crescents above and below each eye and two white wing bars.

Each spring, Northern Parulas arrive mostly from the Caribbean, although they also winter in Mexico, Guatemala, and Belize. Once in the Piedmont, these insectivorous birds move quickly, gleaning leaves and branches high in the canopy, although sometimes they can be seen at eye-level. Since they are difficult to find so high in the trees, North Parulas are often identified by their buzzy, ascending call that end in a distinct, sharp down note. They typically breed in bottomland forests and they make tiny nests of lichen (or Spanish moss further south) high in the canopy of oaks, maples, birches, and sycamores near the tips of branches. Nests are built quickly, in only a few days, and the same nest site may be used year after year. Only two weeks after laying speckled, creamy white eggs, helpless and unseeing young hatch.

Northern Parula populations appear to be stable or even increasing, but an unusual break occurs in their distribution between the north and south. Ornithologists have suggested that this break may be due to habitat change or to air pollution. Air pollution often kills vulnerable lichen species, which are an important nesting material for Northern Parulas.

References:

Cook, D. 2001. The Piedmont Almanac. Raleigh, NC: Barefoot Press.

Daniels, J. C. (2003). Butterflies of the Carolinas. Cambridge, MN: Adventure Publications, Inc.

Ehrlich, P., D. Dobkin, and D. Wheye (1988). The Birder's Handbook. New York: Simon & Schuster.

LeGrand, H. E. Jr. 2009. Notes on the Butterflies of North Carolina. Available at: http://149.168.1.196/nbnc/

LeGrand, H. E. Jr. and Howard, T. E. Jr. 2011. Notes on the Odonates of North Carolina. 3rd Approximation.

Martof, B. S., W. M. Palmer, J. R. Bailey, and J. R. Harrison. 1980. Amphibians and Reptiles of the Carolinas and Virginia. The University of North Carolina Press: Chapel Hill, NC.

Moldenhauer, R. R., and D. J. Regelski. 1996. Northern Parula (Parula americana). In The Birds of North America, No. 215 (A. Poole and F. Gill, eds.). The Academy of Natural Sciences, Philadelphia, PA, and The American Ornithologists' Union, Washington, D.C.